1. Western Australian Herbarium, Department of Parks and Wildlife, Locked Bag 104, Bentley Delivery Centre, Western Australia 6983, Australia; and School of Veterinary and Life Sciences, Murdoch University, Murdoch, Western Australia 6150, Australia
2. Department of Biology, University of New Brunswick, Fredericton, E3B 5A3, New Brunswick, Canada
3. Institut de Systématique, Évolution, Biodiversité, ISYEB – UMR 7205 – CNRS, MNHN, UPMC, EPHE, Muséum national d'Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 39 75005, Paris, France
4. Universitat de Girona, Faculty of Sciences, Department of Environmental Sciences, 17071 Girona, Spain