1. School of Life Sciences, 3190 Maile Way, University of Hawai`i, Honolulu, HI, 96822, USA
2. Western Australian Herbarium, Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, Locked Bag 104, Bentley Delivery Centre, WA 6983, Australia
3. Department of Biology, College of Charleston, 66 George St, Charleston, SC 29424, USA
4. NOAA, Papahānaumokuākea Marine National Monument, 1845 Wasp Boulevard, Building 176, Honolulu, HI 96818, USA