1. College of Fisheries, Tianjin Agricultural University, Tianjin 300384, China
2. Key Laboratory of South China Sea Fishery Resources Exploitation and Utilization, Ministry of Agriculture and Rural Affairs, South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Guangzhou, Guangdong 510300, China
3. Sanya Tropical Fisheries Research Institute, Sanya, Hainan 572019, China
4. Raoping West Coast Biotechnology Co., LTDRaoping, Guangdong 510300, China
5. Key Laboratory of Aquatic Genomics, Ministry of Agriculture and Rural Affairs, Chinese Academy of Fishery Science, Beijing 100141, China