1. Department of Biology, University of Antwerp, Universiteitsplein 1, B-2610 Antwerpen, Belgium
2. UMR 7179 CNRS/MNHN Département d'Ecologie et de Gestion de la Biodiversité, 57 rue Cuvier, Case postale 55, 75231, Paris Cedex 5,France
3. Department of Movement and Sports Sciences, Ghent University, Watersportlaan 2, B-9000 Gent, Belgium