1. Centre for Genomic and Experimental Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh EH4 2XU, UK
2. Disease Models & Mechanisms, The Company of Biologists, Bidder Building, Station Road, Histon, Cambridge CB24 9LF, UK
3. Department of Physiology and Genome Editing Rat Resource Center, Medical College of Wisconsin, Milwaukee, WI 53226, USA