1. College of Agronomy, Qingdao Agricultural University 1 , Qingdao 266109 , China
2. Institute of Crop Sciences, Chinese Academy of Agricultural Sciences (CAAS) 2 , Beijing 100081 , China
3. College of Plant Science and Technology, Huazhong Agricultural University 3 , Wuhan 430070 , China
4. College of Agronomy and Biotechnology, China Agricultural University 4 , 2 Yuanmingyuan West Road, Beijing 100094 , China
5. State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences 5 , Beijing 100101 , China