1. Memorial Sloan Kettering Cancer Center 1 , 1275 York Avenue, New York, NY 10065 , USA
2. Department of Molecular, Cellular, and Developmental Biology 2 , Yale University, New Haven, CT 06511 , USA
3. MRC London Institute of Medical Sciences 3 , Hammersmith Campus, Du Cane Road, London W12 0HS , UK
4. Hammersmith Campus 4 Faculty of Medicine, Imperial College London , , Du Cane Road, London W12 0HS , UK
5. University of Utah School of Medicine and Howard Hughes Medical Institute 5 Department of Biochemistry , , 15 N. Medical Drive East, Salt Lake City, UT 84108 , USA