1. Skirball Institute of Biomolecular Medicine, NYU Grossman School of Medicine 1 , New York, NY 10016 , USA
2. NYU Grossman School of Medicine 2 Department of Cell Biology , , New York, NY 10016 , USA
3. 3 University of Virginia, Department of Biology, 90 Geldard Drive, Physical Life Science Building Room 318, Charlottesville, VA 22904, USA