1. Institute of Evolutionary and Ecological Sciences, Integrative Zoology,Leiden University, POB 9516, 2300 RA Leiden, The Netherlands
2. Department of Experimental Zoology, Wageningen Agricultural University,Marijkeweg 40, 6709 PG Wageningen, The Netherlands
3. Department of Fishculture and Fisheries, Wageningen Agricultural University, Marijkeweg 40, 6709 PG Wageningen, The Netherlands