1. Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, 444-8787, Aichi, Japan
2. The Graduate University of Advanced Studies, Okazaki, 444-8787, Aichi, Japan
3. Department of Embryology, Nara Medical University, Kashihara, 634-0813, Nara, Japan
4. Department of Bioscience, Tokyo University of Agriculture, Setagaya-ku, 156-8502, Tokyo, Japan
5. Division of Stem Cell Therapy, Institute of Medical Science, The University of Tokyo, Minato-ku, 108-8639, Tokyo, Japan
6. Institute for Stem Cell Biology and Regenerative Medicine, Department of Genetics, Stanford University School of Medicine, Stanford, CA, 94305, USA