1. Humphrey Oei Institute of Cancer Research, National Cancer Centre Singapore 1 Division of Cellular & Molecular Research , , Singapore 168583 , Singapore
2. Institute of Molecular & Cellular Biology, Agency for Science, Technology and Research (A*STAR) 2 , Singapore 138673 , Singapore
3. Department of Anatomy, Yong Loo Lin School of Medicine, National University of Singapore 3 , Singapore 117594 , Singapore