1. Center for Stem Cell Biology, Vanderbilt University, Nashville, TN 37232, USA
2. Department of Molecular Physiology and Biophysics, Vanderbilt University, Nashville, TN 37232, USA
3. Department of Cell and Developmental Biology, Vanderbilt University, Nashville, TN 37240, USA
4. Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, PA 19104, USA
5. Institute for Biomedical Informatics, University of Pennsylvania School of Medicine, Philadelphia, PA 19104, USA