1. Trent University 1 Environmental and Life Sciences Graduate Program , , Peterborough, ON , Canada , K9L 0G2
2. Great Lakes Laboratory for Fisheries and Aquatic Sciences, Fisheries and Oceans Canada 2 , Burlington, ON , Canada , L7S 1A1
3. Trent University 3 Department of Biology , , Peterborough, ON , Canada , K9L 0G2