1. Institute of Biological and Environmental Sciences; University of Aberdeen; Tillydrone Avenue Aberdeen AB24 2TZ Scotland UK
2. Marine Laboratory; Marine Scotland - Science; PO Box 101 375 Victoria Road Aberdeen AB11 9DB Scotland UK
3. Institute of Marine Resources; and Ecosystem Studies and Aquaculture and Fisheries Group; IMARES; Haringkade 1 IJmuiden 1976 CP The Netherlands