1. Institute of Ophthalmology, University College London, London EC1V 9EL, UK
2. Division of Biomedical Sciences, Developmental Biomedical Science Laboratory, Graduate School of Biological Sciences, Nara Institute of Science and Technology (NAIST), 8916-5, Takayama-cho, Ikoma 630-0192, Japan
3. Department of Cell and Developmental Biology, University College London, Gower Street, London WC1E 6BT, UK