1. Department of Neuroscience, Developmental Biology Center and Stem Cell Institute, University of Minnesota, Minneapolis, MN 55455, USA
2. Graduate Program in Neuroscience, University of Minnesota, Minneapolis, MN 55455, USA
3. Department of Genetics, Cell and Developmental Biology, Developmental Biology Center and Stem Cell Institute, University of Minnesota, Minneapolis, MN 55455, USA
4. Department of Pharmacology, Graduate School of Medicine, Kyoto University, Kyoto 606-8501, Japan
5. Institute for Protein Research, Osaka University, Osaka 565-0871, Japan
6. Department of Neuroscience, University of Texas Southwestern Medical Center, Dallas, TX 75390, USA