1. Department of Human Anatomy and Psychobiology, University of Murcia, School of Medicine, Murcia, E30071, Spain.
2. Centro de Biología Molecular Severo Ochoa, CSIC-UAM, Madrid, Spain.
3. CIBER de Enfermedades Raras (CIBERER).
4. Department of Cell and Developmental Biology, University College London, Gower Street, London WC1E 6BT, UK.