1. Neural Development Unit, UCL Institute of Child Health, 30 Guilford Street, London WC1N 1EH, UK.
2. Department of Cell and Developmental Biology, UCL, Gower Street, London WC1E 6BT, UK.
3. Howard Hughes Medical Institute, Laboratory of Mammalian Cell Biology & Development, The Rockefeller University, New York, NY 10065, USA.