1. Université Clermont Auvergne, Laboratoire GReD, Clermont-Ferrand F-63000, France
2. Inserm, UMR1103, Clermont-Ferrand F-63001, France
3. CNRS, UMR6293, Clermont-Ferrand F-63001, France
4. Goodman Cancer Research Centre, Department of Human Genetics, McGill University, 1160 Pine Avenue West, rm419, Montreal, Quebec, Canada H3A 1A3