1. Department of Biological Sciences, University of Rhode Island, 120 Flagg Road, Kingston, RI 02881, USA
2. Department of Animal Biology, University of Illinois Urbana-Champaign, Victor E. Shelford Vivarium, Champaign, IL 61820, USA
3. Department of Biological Sciences, University of Alaska Anchorage 3101 Science Circle, Anchorage, AK 99508, USA
4. Department of Ecology and Evolutionary Biology, Brown University, Providence, RI 02912, USA