1. Department of Herpetology, California Academy of Sciences, 825 Howard Street, San Francisco, CA 94013, USA
2. Department of Biological Sciences, California State University, East Bay,Hayward, CA 94542, USA
3. Department of Biology, Portland State University, Portland, OR 97207-0751,USA
4. Zoology, School of Animal Biology MO92, University of Western Australia,Crawley, Western Australia 6009, Australia