1. University of Toronto Mississauga 1 Department of Biology , , Mississauga, ON , Canada , L5L 1C6
2. University of Toronto 2 Department of Ecology and Evolutionary Biology , , Toronto, ON , Canada , M5S 3B2
3. School of Biodiversity, One Health, and Comparative Medicine, College of Medical, Veterinary, and Life Sciences, University of Glasgow 3 , Glasgow, G61 1QH , UK
4. Chongqing Normal University 4 Laboratory of Evolutionary Physiology and Behavior, Chongqing Key Laboratory of Conservation and Utilization of Freshwater Fishes, Animal Biology Key Laboratory of Chongqing Education Commission , , Chongqing 401331 , China
5. UMR Marine Biodiversity, Exploitation, and Conservation, Université Montpellier, CNRS, IRD, IFREMER, INRAE 5 , 34090 Montpellier , France