1. Biology Centre, Czech Academy of Sciences, Institute of Plant Molecular Biology, České Budějovice 37005, Czech Republic
2. Department of Biological Science, College of Bioscience and Biotechnology, Chungnam National University, Daejeon 34134, Republic of Korea
3. Department of Breeding Research, Leibniz Institute of Plant Genetics and Crop Plant Research (IPK) Gatersleben, 06466 Seeland, Germany