1. School of Biological Sciences, University of Reading, Reading, RG6 6AH, U.K.
2. Department of Biology, University of Padova, Padova, Via Ugo Bassi 58/B, 35121, Italy
3. Royal Veterinary College, University of London, London, NW1 0TU, U.K.
4. Department of Neurodegenerative Disease, UCL Queen Square Institute of Neurology, University College London, London, WC1N 3BG, U.K.