1. Graduate School of Life Science, Hokkaido University, Sapporo 060-8589, Japan
2. Graduate School of Agriculture, Hokkaido University, Sapporo 060-8589, Japan
3. Research Faculty of Agriculture, Hokkaido University, Sapporo 060-8589, Japan
4. Faculty of Agriculture, Shizuoka University, Shizuoka 422-8529, Japan