1. Schepens Eye Research Institute, Mass Eye and Ear Infirmary and Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States
2. Schepens Eye Research Institute, Mass Eye and Ear Infirmary and Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States 2University of Exeter Medical School, Exeter, Devon, United Kingdom
3. Department of Cell Biology, Harvard Medical School, Boston, Massachusetts, United States
4. Schepens Eye Research Institute, Mass Eye and Ear Infirmary and Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States 4Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States