1. Department of Cell Biology UCL Institute of Ophthalmology, London, United Kingdom
2. Orbital clinic, Moorfields Eye Hospital and the National Institute for Health Research (NIHR) Biomedical Research Centre at Moorfields Eye Hospital NHS Foundation Trust and UCL Institute of Ophthalmology, London, United Kingdom
3. Department of Cell Biology UCL Institute of Ophthalmology, London, United Kingdom 2Orbital clinic, Moorfields Eye Hospital and the National Institute for Health Research (NIHR) Biomedical Research Centre at Moorfields Eye Hospital NHS Foundation Trust and