1. Department of Biochemistry, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
2. nstitute for Aging and Metabolic Diseases, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
3. Department of Biomedicine & Health Sciences, Graduate School, College of Medicine, The Catholic University of Korea, Seoul 06591, Korea
4. These authors contributed equally to this work.
5. Genome Editing Research Center, Korea Research Institute of Bioscience and Biotechnology (KRIBB), Daejeon 34141, Korea