1. Department of Genome Medicine, Mie University Graduate School of Medicine, Tsu, Mie 514‑8507, Japan
2. Department of Gastrointestinal and Pediatric Surgery, Division of Reparative Medicine, Institute of Life Sciences, Mie University Graduate School of Medicine, Tsu, Mie 514‑8507, Japan
3. California Institute of Technology, Pasadena, CA 91125, USA
4. Department of Genome Medicine, Division of Reparative Medicine, Institute of Life Sciences, Mie University Graduate School of Medicine, Tsu, Mie 514‑8507, Japan