1. Agricultural Biotechnology Department, National Institute of Agricultural Sciences, Rural Development Administration, 370 Nongsaengmyeong-ro, Jeonju-si, Jeollabuk-do, 54874, Korea
2. Maize Experiment Station, Gangwondo Agricultural Research and Extension Services, Hongcheon, 25160, Korea
3. Crop Foundation Research division, National Institute of Crop Science, Rural Development Administration, 181 Hyeoksin-ro, Iseo-myeon, Wanju-Gun, Jeollabuk-do, Korea
4. Graduate School of International Agricultural Technology and Crop Biotechnology Institute/GreenBio Science & Technology, Seoul National University, Pyeongchang 25354, Korea