1. Department of Wood Science , University of British Columbia , Vancouver , BC V6T 1Z4 , Canada
2. Forestry Division , York Timbers , Sabie , Mpumalanga , South Africa
3. Institute for Commercial Forestry Research , Pietermaritzburg , KZN , South Africa
4. CharlieC Consulting , Cheltenham , Gloucestershire , GL54 4NU , UK
5. Department of Botany , University of British Columbia , Vancouver , BC V6T 1Z4 , Canada