1. Flower Research and Development Centre, Zhejiang Academy of Agricultural Sciences, Hangzhou311202, China
2. Key Laboratory of Crop Ecophysiology and Farming Systems in the Middle Reaches of the Yangtze River, Ministry of Agriculture, College of Plant Science and Technology, Huazhong Agricultural University, Wuhan430070, China
3. School of Agricultural Science, Yunnan University, Kunming650091, China