1. Animal Resource Science & Management, Faculty of Resource Science and Technology, Universiti Malaysia Sarawak , 94300 Kota Samarahan , Sarawak , Malaysia
2. Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak , 94300 Kota Samarahan , Sarawak , Malaysia
3. Wildlife Biology Program and Division of Biological Sciences, University of Montana , Missoula MT 59812, USA