1. Departments of Zoology & Botany; University of British Columbia; 3529-6270 University Boulevard Vancouver British Columbia V6T 1Z4 Canada
2. Biodiversity Research Centre; University of British Columbia; 3529-6270 University Boulevard Vancouver British Columbia Canada
3. Department of Natural Resources Management, Box 42125, Goddard Hall; Texas Tech University; Lubbock TX 79409 U.S.A.
4. Department of Zoology; Michigan State University; East Lansing MI 48824 U.S.A.
5. Institute for Tropical Biology and Conservation; Universiti Malaysia Sabah, Jalan UMS; 88400 Kota Kinabalu Sabah Malaysia
6. Department of Zoology, Faculty of Resource Science and Technology; Universiti Malaysia Sarawak; 94300 Kota Samarahan Sarawak Malaysia
7. Sabah Wildlife Department; Ibu Pejabat Tingkat 5, Blok B Wisma Muis 88100 Kota Kinabalu Sabah Malaysia