1. Ethiopian Institute of Agricultural Research, Pawe Agricultural Research Center, Pawe, Ethiopia
2. CIMMYT-Ethiopia, ILRI Campus, CMC Road, P.O. Box 5689, Addis Ababa, Ethiopia.
3. Department of Microbial, Cellular and Molecular Biology, Addis Ababa University, College of Natural Sciences, Addis Ababa, Ethiopia
4. Tennessee State University, Nashville, USA