1. Department of Biological and Environmental Sciences, Walter Sisulu University, Mthatha, South Africa
2. Department of Ichthyology and Fisheries Science, Rhodes University, Makanda, South Africa
3. Alfred-Wegener-Institute, Helmholtz Center for Polar and Marine Research, Bremerhaven, Germany
4. South African Institute for Aquatic Biodiversity, Makhanda, South Africa
5. Department of Zoology and Entomology, University of Fort Hare, Alice, South Africa