1. Chiba Prefectural Agriculture and Forestry Research Center, Chiba, Japan
2. Institute of Agrobiological Sciences, National Agriculture and Food Research Organization, Ibaraki, Japan
3. Department of Life Sciences, Toyo University, Gunma, Japan
4. Institute of Crop Science, National Agriculture and Food Research Organization, Ibaraki, Japan
5. Institute of Vegetable and Floriculture Science, National Agriculture and Food Research Organization, Ibaraki, Japan