1. Department of Veterinary Medicine, Guangdong Ocean University, Zhanjiang, China;
2. Shenzhen Institute of Guangdong Ocean University, Shenzhen, China;
3. Department of Animal Science, Guangdong Ocean University, Zhanjiang, China;
4. Faculty of Agriculture and Life Sciences, Lincoln University, Lincoln, New Zealand;
5. Department of Biomedical and Diagnostic Sciences College of Veterinary Medicine, The University of Tennessee, Knoxville, TN, USA;
6. MOA Key Laboratory for Animal Vaccine Development Key Laboratory of Zoonoses Control and Prevention of Guangdong College of Veterinary Medicine, South China Agricultural University, Guangzhou, China