1. a Key Laboratory of Biology and Genetic Resources of Tropical Crops, Ministry of Agriculture, Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences , Haikou , 571101 , China
2. b Key Laboratory of Protection and Developmental Utilization of Tropical Crop Germplasm Resources, Ministry of Education , Haikou , 570228 , China
3. c College of Horticulture and Landscape Architecture, Hainan University , Haikou , 570228 , China