1. Key Laboratory of Tibetan Medicine Research, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, China
2. University of Chinese Academy of Sciences, Beijing 100039, China
3. Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining 810016, China
4. CAS Key Laboratory of Microbial Physiological and Metabolic Engineering, Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China