1. Laboratory of Aromatic and Medicinal Plants, Center of Biotechnology of Borj Cedria, Hammam Lif, BP, Tunisia
2. Laboratory of Bioactive Substances, Center of Biotechnology of Borj Cedria, Hammam Lif, BP, Tunisia
3. Department of Biology, College of Science, University of Hai’l, Hai’l, Saudi Arabia
4. Laboratory of Genetics, Biodiversity and Valorisation of Bioresources, High Institute of Biotechnology, University of Monastir 5000, Tunisia