1. Department of Systematic Zoology and Ecology, Eötvös Loránd University, 1117 Budapest, Pázmány Péter sétány 1/C, Hungary
2. Institute of Landscape Ecology, Münster University, Heisenbergstraße 2, 48149 Münster, Germany
3. Department of Anatomy, Cell- and Developmental Biology, Eötvös Loránd University, 1117 Budapest, Pázmány Péter sétány 1/C, Hungary