1. Department of Systematic Zoology and Ecology, Eötvös Loránd University, 1117 Budapest, Pázmány Péter sétány 1/C, Hungar
2. Department of Anatomy, Cell- and Developmental Biology, Eötvös Loránd University, 1117 Budapest, Pázmány Péter sétány 1/C, Hungary
3. University of Münster, Institute of Landscape Ecology, Heisenbergstraße 2, 48149 Münster, Germany