1. National Institute of Marine Sciences and Technologies (INSTM), Salammbô, Tunis, Tunisia
2. University of Tunis El Manar, Faculty of Sciences of Tunis, Laboratory of Diversity, Management and Conservation of Biological Systems (LR18ES06), Tunis, Tunisia
3. University of Poitiers, Laboratory of Ecology and Biology of Interactions (UMR CNRS 7267 EBI), Ecology Evolution Symbiosis Team, Poitiers Cedex 9, France