1. Biology Department, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran;
2. Institute of Applied Zoology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran;
3. Department of Biology, Faculty of Sciences, Golestan University, Gorgan, Iran;
4. Department of Biotechnology and Plant Breeding College of Agriculture, Ferdowsi University of Mashhad, Mashhad, Iran