1. CAS Key Laboratory of Biobased Materials, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, P. R. China
2. College of Life Science, Qingdao University, Qingdao, P. R. China
3. Key Laboratory for Tobacco Gene Resources, Tobacco Research Institute, Chinese Academy of Agricultural Sciences, Qingdao, P. R. China
4. School of Biological Science, Jining Medical University, Jining, P. R. China