1. Department of Medicine, Addenbrooke’s Hospital, University of Cambridge, Cambridge, UK;
2. Parasites and Microbes, Wellcome Sanger Institute, Hinxton, UK;
3. Department of Archaeology, University of Cambridge, Cambridge, UK;
4. Centre for Microbial Ecology and Genomics, Department of Biochemistry, Genetics and Microbiology, University of Pretoria, Hatfield, South Africa;
5. Human Origins and Palaeo-Environments Research Group, Department of Anthropology and Geography, Oxford Brookes University, Oxford, UK