1. Faculty of Sciences, Department of Environmental Sciences, University of Girona, Ma Aurèlia Capmany 69, Girona 17003, Spain
2. Phycology Research Group and Centre for Molecular Phylogenetics and Evolution, Ghent University, Krijgslaan 281, Building S8, Ghent 9000, Belgium
3. Department of Biodiversity, Conservation and Attractions, Western Australian Herbarium, Biodiversity and Conservation Science, Kensington, Locked Bag 104, Bentley Delivery Centre WA 6983, Australia
4. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, R.O.C.