1. Department of Marine Science, School of Natural Sciences, Incheon National 119 Academy-ro, Yeonsu-gu, Incheon 22012, Republic of Korea
2. Research Institute of Basic Sciences, Incheon National University, Incheon 22012, Republic of Korea
3. Department of Ecology and Evolutionary Biology, University of Connecticut, Stamford, Connecticut 06901, USA
4. Department of Natural Sciences, University of Alaska Southeast, 11120 Glacier Highway, Juneau, Alaska 99801, USA