1. Ifremer, LITTORAL, 29900 Concarneau, France
2. Ecosystem-Based Management of Marine Resources Program, Environment & Life Sciences Research Center, Kuwait Institute for Scientific Research, P.O. Box 1638, Salmiya, 22017, Kuwait
3. Université de Bretagne Occidentale, Station de Biologie Marine, Place de la Croix, F-29900 Concarneau, France
4. Senckenberg am Meer, German Center for Marine Biodiversity Research (DZMB), Südstrand 44, D-26382 Wilhelmshaven, Germany